RAWALPINDI, Pakistan -- The stunning arrest of Sept. 11 mastermind Khalid Shaikh Mohammed by Pakistani intelligence and CIA agents began with a near miss last month in a dusty border town and became a feverish chase fueled by communications intercepts and suspects' interrogations, security officials said Sunday.

Mohammed, an unidentified man of Middle Eastern origin and Pakistani Ahmed Abdul Qadus were arrested early Saturday in Rawalpindi, a bustling city adjacent to the capital. Mohammed and the unidentified man were handed over to U.S. authorities and spirited to an undisclosed location out of the country, a senior government official told The Associated Press on condition of anonymity.

Security officials, also speaking on condition they not be named, said the trail heated up after authorities arrested an Egyptian man during a raid in the frontier city of Quetta on Feb. 14. Authorities had hoped to find Mohammed -- a top al-Qaida figure with links to a decade of deadly plots -- but he was not there.

"At the time of that raid in Quetta the authorities were looking for Khalid Shaikh but he escaped and from there they followed him to Rawalpindi," the senior government official said. "They got information from the man they picked up in Quetta and from phone calls until they tracked him down to Rawalpindi."

Changed story

A top police official in Quetta said the arrested suspect changed his story many times during questioning, but finally identified himself as Abdul Rehman from Egypt. The official said police were aware that another suspect got away, but were not told of his significance. The police later handed Rehman over to Pakistan's intelligence agency, known as ISI.

"We got some information about two foreigners who were in the neighborhood. When we went there we found only one," the official said. "Rehman admitted there was someone else with him, but he never said anything about Khalid Shaikh Mohammed."

The second foreign suspect arrested in Rawalpindi with Mohammed has not been named, but the senior government official said he was "proving to be an important man."

Mohammed, 37, is perhaps the most senior al-Qaida member after bin Laden and his deputy, Ayman al-Zawahri. He allegedly organized the Sept. 11 terror attacks, and was linked to a 1995 plot to bomb trans-Pacific airliners and crash a plane into CIA headquarters. He also has been tied to April's bombing of a synagogue in Tunisia.

A Pakistani intelligence official with knowledge of the case said American communications experts helped Pakistani authorities trace an e-mail the arrested Quetta suspect sent to Qadus. They immediately put him under surveillance, which led authorities to Mohammed.

U.S. officials said the operation in which Mohammed was captured was carried out jointly by the CIA and Pakistani officers.

Qadus' family on Sunday vehemently denied his involvement in terrorism and insisted he was alone at home with his wife and children when authorities burst in around 3 a.m. Saturday.

Agents, all of whom appeared to be Pakistani, took a computer hard-drive, documents and U.S. dollars from the house, the family and security officials said. Khanum said the computer had no Internet hookup and that her brother didn't even know how to use it.

"He is slow, and he is so innocent and friendly, it is inconceivable that he could be involved in intrigue. People run rings around him because it would never occur to him that they might lie or take advantage of him," she said from the family's living room in an upscale neighborhood of the city.

Qadus' father, Abdul Qadus, is a prominent microbiologist who worked in Africa for the World Health Organization for many years before retiring, Khanum said. The 42-year-old Qadus, however, has never been able to hold down a job and had lived at home with his parents his entire life, she said.

Qadus' mother, Mahlaqa, is a local leader of Jamaat-e-Islami, one of the main hardline religious parties in Pakistan. The family speculated the arrest was a political ploy to discredit her and the party, which is part of an ultraconservative coalition that came in third in last year's parliamentary elections, largely on the strength of a virulently anti-American platform.

Ameer-ul Azeem, spokesman for Jamaat-e-Islami, said Qadus had not done anything wrong.

"Arabs, Afghans, Sudanese, it is not a sin to host them as guests, unless their crime is proven," he said when asked whether Qadus or his family had links with al-Qaida.

Interior Ministry spokesman Iftikhar Ahmed brushed off the family's claims of innocence, saying Qadus, Mohammed and the third man were all arrested at the family's home.

"There was a single operation," Ahmed said. "Naturally, they will deny it. Everybody says they are innocent, and you can draw your own conclusion."

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AP Correspondent Munir Ahmad contributed to this report)